Fossil range: Cambrian (or earlier?) - present
|Subphyla & Classes|
† = extinct
Echinoderms (Phylum Echinodermata) are a phylum of marine animals (including sea stars) found at all ocean depths. Brittle stars, or ophiurids, are Echinoderms closely related to Sea stars. Brain coral is a Common name given to several genera of Coral which are characterized by the Spheroid shape of their colonies, and Eumetazoa is a Clade comprising all major Animal groups except sponges Characteristics of eumetazoans include true tissues organized into Deuterostomes (taxonomic term Deuterostomia; from the Greek "second mouth" are a superphylum of Animals They are a subtaxon of the Jacob Theodor Klein nickname Plinius Gedanensium ( 15 August 1685 &ndash 27 February 1759) was a Royal Prussian Jurist The Stylophora are a class within the Subphylum Homalozoa of the Phylum Echinodermata. Crinoids, also known as sea lilies or feather-stars, are marine animals that make up the class Crinoidea of the Echinoderms (phylum Echinodermata Brittle stars, or ophiurids, are Echinoderms closely related to Sea stars. Starfish (also called sea stars) are any Echinoderms belonging to the class Asteroidea. Sea urchins are small globular spiny sea cat animals composing most of class Echinoidea. The sea cucumber (also known as trepang, bêche-de-mer, or ambiguously Sea slug) is an Echinoderm of the class Holothuroidea Helicoplacus (often misspelled Helioplacus) is the earliest well-studied Fossil echinoderm. Arkarua is a small Precambrian disk-like Fossil with a raised center a number of radial ridges on the rim and a five-pointed central depression marked Cothurnocystis along with all of its relatives in Stylophora, are enigmatic organisms whose relationship with other organisms namely the Echinodermata The Edrioasteroids are an extinct class of echinoderm that lived from the Ediacaran (if Arkarua was indeed an edrioaster to Blastoids (class Blastoidea are an Extinct type of stemmed Echinoderm. Blastoids (class Blastoidea are an Extinct type of stemmed Echinoderm. The Eocrinoidea ("dawn crinoids" are a paraphyletic division of Echinoderms in that the Blastoids are derived/descended from them Otto Max Johannes Jaekel ( February 21, 1863 &ndash March 6, 1929) was a German Paleontologist and Geologist A phylum ( Plural: phyla) is a Taxonomic rank between Kingdom and above Class. Marine is an Umbrella term. As an adjective it is usually applicable to things relating to the Sea or Ocean, such as Marine biology, Marine Starfish (also called sea stars) are any Echinoderms belonging to the class Asteroidea. Aside from the problematic Arkarua, the first definitive members of the phylum appeared near the start of the Cambrian period, and contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates; they are the largest phylum without freshwater or terrestrial representatives. Arkarua is a small Precambrian disk-like Fossil with a raised center a number of radial ridges on the rim and a five-pointed central depression marked The Cambrian is a geologic period and system that began about Ma (million years ago at the end of the Proterozoic eon and ended about Ma with In Biology, a species is one of the basic units of Biological classification and a Taxonomic rank. Deuterostomes (taxonomic term Deuterostomia; from the Greek "second mouth" are a superphylum of Animals They are a subtaxon of the Chordates ( Phylum Chordata) are a group of Animals that includes the Vertebrates together with several closely related Invertebrates The word derives from the Greek εχινοδέρματα (echinodermata), plural of εχινόδερμα (echinoderma), "spiny skin" and that from εχινός (echinos), "sea-urchin", originally "hedgehog" + δέρμα (derma), "skin". Greek (el ελληνική γλώσσα or simply el ελληνικά — "Hellenic" is an Indo-European language, spoken today by 15-22 million people mainly
The Echinoderms are important both biologically and geologically: biologically because few other groupings are so abundant in the biotic desert of the deep sea, as well as the shallower oceans, and geologically as their ossified skeletons are major contributors to many limestone formations, and can provide valuable clues as to the geological environment. Further, it is held by some that the radiation of echinoderms was responsible for the Mesozoic revolution of marine life. The Mesozoic Era is one of three geologic eras of the Phanerozoic eon.
Two main subdivisions of Echinoderms are traditionally recognised: the more familiar, motile Eleutherozoa, which encompasses the Asteroidea (starfish), Ophiuroidea (brittle stars), Echinoidea (sea urchin and sand dollar) and Holothuroidea (sea cucumbers); and the sessile Pelmatazoa, which consist of the crinoids. Eleutherozoa is a Subphylum of Echinoderms They are mobile animals with the mouth diceted towards the substrate Starfish (also called sea stars) are any Echinoderms belonging to the class Asteroidea. Brittle stars, or ophiurids, are Echinoderms closely related to Sea stars. Sea urchins are small globular spiny sea cat animals composing most of class Echinoidea. Sand dollars (order Clypeasteroida are flat round marine animals related to sea urchins ( Echinoids, sea stars and other echinoderms The sea cucumber (also known as trepang, bêche-de-mer, or ambiguously Sea slug) is an Echinoderm of the class Holothuroidea Crinoids, also known as sea lilies or feather-stars, are marine animals that make up the class Crinoidea of the Echinoderms (phylum Echinodermata Some crinoids, the feather stars, have secondarily re-evolved a free-living lifestyle. Crinoids, also known as sea lilies or feather-stars, are marine animals that make up the class Crinoidea of the Echinoderms (phylum Echinodermata
A fifth class of Eleutherozoa consisting of just two species, the Concentricycloidea (sea daisies), were recently merged into the Asteroidea. Sea daisies (Infraclass Concentricycloidea order Peripoda) make up an unusual group of deep-sea taxa belonging to the phylum Echinodermata, with three The fossil record contains a host of other classes which do not appear to fall into any extant crown group. The Crown Group is a Catering and event caterer in the United Kingdom.
Echinoderms evolved from animals with bilateral symmetry; although adult echinoderms possess radial symmetry. "Bilateral symmetry" redirects here For bilateral symmetry in mathematics see Reflection symmetry. "Bilateral symmetry" redirects here For bilateral symmetry in mathematics see Reflection symmetry. Echinoderms' larvae are ciliated, free-swimming organisms that organize in a bilaterally symmetric fashion that makes them look like embryonic chordates. A cilium (plural cilia) is an Organelle found in eukaryotic cells Cilia are tail-like projections extending approximately Chordates ( Phylum Chordata) are a group of Animals that includes the Vertebrates together with several closely related Invertebrates Later, the left side of the body grows at the expense of the right side, which is eventually absorbed. The left side then grows in a pentaradially symmetric fashion, in which the body is arranged in five parts around a central axis. "Bilateral symmetry" redirects here For bilateral symmetry in mathematics see Reflection symmetry.
All echinoderms exhibit fivefold radial symmetry in portions of their body at some stage of life, even if they have secondary bilateral symmetry. Many crinoids and some starfish exhibit symmetry in multiples of the basic five, with starfish such as Helicoilaster spp. known to possess up to 50 arms, and the sea-lily Comanthina schlegelii boasting 200.
With a radially symmetrical animal, the traditional designation of a front and back, posterior and anterior becomes troublesome — what does one term the front? To get around this, a different terminology is used with the Echinoderms. The mouth and anus of the developing adult migrate simultaneously from the front and back of the larvum to opposite ends of the organism, allowing the description of an "oral" (containing the mouth) and opposite "aboral" side. Grazing organisms such as sea urchins tend to have their mouth on the substrate upon which they are feeding, and their anus on their "top" surface; burrowers such as sea cucumbers have their mouth at their front, and the anus behind their direction of travel. Hence the sea-cucumbers appear to have evolved from a sea-urchin-like organism which gradually tipped on its side and lengthened.
The individual limbs can be named on the basis of a break in the symmetry provided by the "filter plate". This lies beside the anus on the aboral plate, between two of the radial arms; the radial limbs are designated letters a to e in a clockwise direction from this marker.
In spite of their potentially misleading name and sometimes foreboding appearance, the echinoderms do not possess an external skeleton. Rather, a thin outermost skin covers a mesodermal endoskeleton made of tiny calcified plates and spines, which forms a rigid support contained within tissues of the organism. A germ layer is a collection of cells formed during animal Embryogenesis. Calcium carbonate is a Chemical compound with the Chemical formula Ca[[Carbon C]] O 3 Some groups, such as the sea urchins, also possess calcareous spines that serve to protect the organism from predation and colonisation by encrusting organisms; the sea cucumbers secondarily use these spines for locomotion. These spines too are covered by a thin layer of epidermis.
The calcite grown by the organisms is diagnostically rich in the element magnesium; they may consist of 3 to 15 % magnesium oxide. The abundance of this small element property confers them a higher skeletal density, and the chemical properties of magnesium encourage it to form stronger bonds — making for a stronger, more resistant skeleton. The feeding apparatus of the echinoderms is particularly enriched in magnesium; the rock-grazing lifestyle of the sea urchins makes their mandibles especially prone to wear, thus the extra strength provides a significant advantage, outweighing the metabolic costs involved in concentrating the magnesium.
Despite the robustness of the individual skeletal modules, complete echinoderm skeletons are rare in the fossil record. This is because they quickly disarticulate once the encompassing skin rots away, and in the absence of tissue there is nothing to hold the plates together. The modular construction is a result of the growth system employed by echinoderms, which adds new segments at the centre of the radial limbs, pushing the existing plates outwards in the fashion of a conveyor belt. The spines of sea urchins are most readily lost, as they are not even attached to the main skeleton in life. Each spine can be moved individually and is thus only loosely attached in life; a walk above a rocky shore will often reveal a large number of spineless but otherwise complete sea urchin skeletons.
Skeletal elements are also deployed in some specialised ways; as well as the famous feeding organ of the sea urchins, the "Aristotle's lantern", crinoids' stalks and the supportive "lime ring" of sea cucumbers consist of specialised calcite plates. Sea urchins are small globular spiny sea cat animals composing most of class Echinoidea.
The epidermis itself consists of cells responsible for the support and maintenance of the skeleton, as well as pigment cells, mechanoreceptor cells, which detect motion on the animal's surface, and sometimes gland cells which secrete sticky fluids or even toxins.
The varied and often vivid colours of the echinoderms are produced by the action of the skin pigment cells. These may be light sensitive, and as a result many species change appearance completely as night falls. The reaction can happen very quickly — the sea urchin Centrostephanus discolours longispinus changes from jet black to grey-brown in just 50 minutes when exposed to light. The colours are produced by a variable combination of coloured pigments, such as the dark Melanin, red Carotinoids, and Carotinproteins, which can be blue, green or violet.
Echinoderms possess a unique water vascular system, a network of fluid-filled canals that function in gas exchange, feeding, and secondarily in locomotion. The water vascular system is a hydraulic system used by Echinoderms such as Starfish and Sea urchins for locomotion food and waste transportation and This system may have allowed them to function without the gill slits found in other Deuterostomes. The system comprises a central ring, the hydrocoel, and radial ambulacra stretching along each limb of the organism. As well as assisting with the distribution of nutrients through the animal, the system is most obviously expressed in the "tube-feet" of most echinoderms. These are extensions of the water vascular system which poke through holes in the skeleton and can be extended or contracted by the redistribution of fluid between the foot and internal sac. In the crinoids, these tube feet waft food particles captured on the radial limbs towards the central mouth; in the asteroids, the same wafting motion is employed to move the animal across the ground. Sea urchins use their feet to prevent the larvae of encrusting organisms from settling on their surfaces; potential settlers are moved to the urchin's mouth and eaten. Some burrowing sea stars poke their tube feet through the surface of the sand or mud above them into the water column and use them to attain oxygen from the water column.
Although echinoderms possess a complete digestive tube (tubular gut), it is very simple, often simply leading directly from mouth to anus. It can generally be divided into a throat, stomach, intestine and rectum. They also possess an open and reduced circulatory system — consisting of a central ring and five radial vessels, but no heart. This is an article about the rock music band "Circulatory System"
They have a simple radial nervous system that consists of a modified nerve net — interconnected neurons with no central brain (although some do possess ganglia. The nervous system is a Network of specialized cells that communicate information about an animal's surroundings and itself For the album by Brian Eno, see Nerve Net (album. A nerve net is a type of simple Nervous system that is found in members of the phylum The brain is the center of the Nervous system in animals All Vertebrates and the majority of Invertebrates have a brain In Anatomy, a ganglion (pl ganglia) is a tissue mass. Neurology In neurological contexts ganglia are composed mainly of ) Nerves radiate from central rings around the mouth into each arm; the branches of these nerves coordinate the movements of the organism.
The gonads of the organisms occupy the entire body cavities of sea urchins and sea cucumbers; the less voluminous crinoids, brittle stars and starfish having two gonads per arm. The gonad is the organ that makes Gametes The gonads in males are the Testes and the gonads in Females are the Ovaries. Whilst the primitive condition is considered to be one genital aperture, many organisms have multiple holes through which eggs or sperm may be released.
Echinoderms become sexually mature after approximately two to three years, depending on the species and the environmental conditions. The eggs and sperm cells are released into open water, where fertilization takes place. The release of sperm and eggs is co-ordinated temporally in some species, and spatially in others. Internal fertilization has currently been observed in three species of starfish, three brittle stars and a deep water sea cucumber.
In some species of feather star, the embryos develop in special breeding bags, where the eggs are held until sperm released by a male happen to find them and fertilize the contents. This can also be found among sea urchins and sea cucumbers, where exhibit care for their young can occur, for instance in a few species of sand dollars who carry their young between the pricks of their oral side, and heart urchins possess breeding chambers. With brittle stars, special chambers can be developed near the stomach bags, in which the development of the young takes place. Species of sea cucumbers with specialized care for their offspring may also nurse the young in body cavities or on their surfaces. In rare cases, direct development without passing through a bilateral larval stage can occur in some starfish and brittle stars. Another strategy that has evolved in some starfish and brittle stars is the ability to reproduce asexually by dividing in two halves while they are small juveniles, while turning to sexual reproduction when they have reached sexual maturity. These species have six arms.
The development of an echinoderm begins with a bilaterally symmetrical embryo, with a coeloblastula developing first. Gastrulation marks the opening of the "second mouth" that places them within the deuterostomes, and the mesoderm, which will host the skeleton, migrates inwards. Gastrulation is a phase early in the development of animal Embryos during which the morphology of the embryo is dramatically restructured by Cell migration. The secondary body cavity, the coelom, forms by the partitioning of three body cavities.
Upon metamorphosis, each taxon produces a distinct larvum, the left hand side of which develops into the adult organism, the right hand side eventually being absorbed; the left hand side typically becomes the oral plate.
F. M. Balfour and D. I. Williamson hold that no echinoderms acquired larvae until after the classes of the phylum were established, i. e. after the Ordovician. The Ordovician is a geologic period and system, the second of six of the Paleozoic era, and covers the time between 488 Some modern brittle stars and heart urchins have no larvae, and they develop as protostomes. Brittle stars, or ophiurids, are Echinoderms closely related to Sea stars. Protostomia (from the Greek: mouth first) are a Taxon of Animals Together with the Deuterostomes and a few smaller phyla H. B. Fell and Williamson (2003) argue that the original echinoderms were radial protostomes and the bilateral larvae were later additions.
Many echinoderms have remarkable powers of regeneration. In Biology, an Organism is said to regenerate a lost or damaged part if the part regrows so that the original function is restored Some sea stars are capable of regenerating lost arms. In some cases, lost arms have been observed to regenerate a second complete sea star. Sea cucumbers often discharge parts of their internal organs if they perceive danger. The discharged organs and tissues are quickly regenerated. Sea urchins are constantly losing their spines through damage — all parts are replaceable. Some starfish populations can reproduce entirely asexually purely by the shedding of arms for long periods of time.
Echinoderms are globally distributed in almost all depths, latitudes and environments in the ocean. They reach highest diversity in reef environments but are also widespread on shallow shores, around the poles — refugia where crinoids are at their most abundant — and throughout the deep ocean, where bottom-dwelling and burrowing sea cucumbers are common — sometimes accounting for up to 90 % of organisms. Whilst almost all echinoderms are benthic — that is, they live on the sea floor — some sea-lilies can swim at great velocity for brief periods of time, and a few deep-sea sea cucumbers are fully floating. Some crinoids are pseudo-planktonic, attaching themselves to floating logs and debris, although this behaviour was exercised most extensively in the Paleozoic, before competition from such organisms as barnacles restricted the extent of the behaviour. Some sea cucumbers employ a similar strategy, hitching lifts by attaching to the sides of fish.
The larvæ of many echinoderms, especially starfish and sea urchins, are pelagic, and with the aid of ocean currents can swim great distances, reinforcing the global distribution of the phylum.
The modes of feeding vary greatly between the constituent taxa. Crinoids and some brittle stars tend to be passive filter-feeders, absorbing suspended particles from passing water; sea urchins are grazers, sea cucumbers deposit feeders, and starfish active hunters.
Crinoids employ a large net-like structure to sieve water as it is swept by currents, and to adsorb any particles of matter sinking from the ocean overhead. Once a particle touches the arms of the creature, the tube feet act to swish it to the central mouth of the crinoid, where it is ingested, nutrients removed, and the remains egested through its anus to the underlying water column.
Many sea urchins graze on the surfaces of rocks, scraping off the thin layer of algae covering the surfaces. Other toothless breeds devour smaller organisms, which they may catch with their tube feet, whole. Sand dollars may perform suspension feeding.
Sea cucumbers may be suspension feeders, sucking vast quantities of sea water through their guts and absorbing any useful matter. Others use their feeding apparatus to actively capture food from the sea floor. Yet others deploy their feeding apparatus as a net, in which smaller organisms become ensnared.
Whilst some starfish are detritovores, extracting the organic material from mud, and others mimic the crinoids' filter feeding, most are active hunters, attacking other starfish or shellfish. The latter are seized and held by the tube feet; starfish then stiffen their legs, expanding the shell. The starfish can use catch connective tissue to lock their arms in place and maintain a force on the prey whilst exerting minimal effort; the unfortunate victim must expend energy resisting the force with its adductor muscle. When the abductor tires, the starfish can insert its stomach through the opening and release gastric juices, digesting the prey alive.
Despite their low nutrition value and the abundance of indigestable calcite, many organisms, such as Crabs, sharks, sea birds and larger starfish, make a living by feeding on echinoderms. Crabs are decapod Crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (βραχύ / brachy Sharks ( Superorder Selachimorpha) are a type of Fish with a full cartilaginous Skeleton and a highly streamlined body Defensive strategies employed include the presence of spines, toxins, which can be inherent or delivered through the tube feet, and the discharge of sticky entangling threads by sea cucumbers. Being stabbed by a sea urchin may result in painful death.
Echinoderms provide a key ecological role in ecosystems. For example, the grazing of sea urchins reduces the rate of colonization of bare rock; the burrowing of sand dollars and sea cucumbers depleted the sea floor of nutrients and encouraged deeper penetration of the sea floor, increasing the depth to which oxygenation occurs and allowing a more complex ecological tiering to develop. Starfish and brittle stars prevent the growth of algal mats on coral reefs, which would obstruct the filter-feeding constituent organisms. Some sea urchins can bore into solid rock; this bioerosion can destabilise rock faces and release nutrients into the ocean. Bioerosion describes the Erosion of hard ocean substrates by living organisms by a number of mechanisms
The echinoderms are also the staple diet of many organisms, most notably the otter; conversely, many sea cucumbers provide a habitat for parasites, including crabs, worms and snails. Otters are semi- aquatic (or in one case aquatic) fish-eating Mammals The otter subfamily Lutrinae forms part of the family The extinction of large quantities of echinoderms appears to have caused a subsequent overrunning of ecosystems by seaweed, or the destruction of an entire reef.
The first universally accepted echinoderms appear in the Cambrian period around . The Cambrian is a geologic period and system that began about Ma (million years ago at the end of the Proterozoic eon and ended about Ma with Echinoderms left behind an extensive fossil record. Despite this, there are numerous conflicting hypotheses on their phylogeny. Based on their bilateral larvae, many zoologists argue that echinoderm ancestors were bilateral and that their coelom had three pairs of spaces (trimeric).
Some have proposed that radial symmetry arose in a free-moving echinoderm ancestor and that sessile groups were derived several times independently from free-moving ancestors. Unfortunately, this view does not address the significance of radial symmetry as an adaptation for a sessile existence.
The more traditional view is that the first echinoderms were sessile, became radial as an adaptation to that existence, and then gave rise to free-moving groups. This view perceives the evolution of endoskeletal plates with stereom structure and of external ciliary grooves for feeding as early echinoderm developments.
The extinct members of Class Homalozoa, commonly referred to as carpoids, had stereom ossicles but were not radially symmetrical, and the status of their water-vascular system is not known. Further, extinct members of the Class Helicoplacoidea possessed three, true ambulacral grooves, and their mouth was on the side of their body.
Attachment to a substratum would have selected for radial symmetry and may have marked the origin of the Class Crinoidea. Members of Crinoidea, along with the extinct members of Class Cystoidea, were primitively attached to a substratum by an aboral stalk. An ancestor that became free-moving might have given rise to Asteroidea, Ophiuroidia, Holothuroidea, and Echinoidea.
Most humans know the Echinoderms rather from the unpleasant side: if one finds oneself near the coast, on a rocky shore or reef, one must beware the poisonous prick of a sea urchin. Because of their fine, circular structure they often break off when removed, and need patience — or the physician — to fully remove. But in the kitchens of some countries, Echinoderms are regarded as a delicacy; and for children sea-urchin skeletons are as popular a collecting object as brightly coloured starfish are fascinating.
The economic impact of Echinoderms is primarily local. Around 50,000 tons of sea urchins are captured each year, the gonads of which are consumed particularly in Japan, Peru and in France. The gonad is the organ that makes Gametes The gonads in males are the Testes and the gonads in Females are the Ovaries. The taste is described as soft and melting, like a mix of seafood and fruit. The quality depends on the color, which can range from light yellow to bright orange.
Sea cucumbers are also considered a delicacy in some countries of south east Asia; particularly popular are the pineapple roller Thelenota ananas (susuhan) and the red Halodeima edulis. Pineapple ( Ananas comosus) is the common name for an edible Tropical Plant and also its Fruit It is native to the southern part of Brazil They are well known as bêche de mer or Trepang in China and Indonesia. The sea cucumbers are dried, and the potentially poisonous entrails removed. The strong poisons of the sea cucumbers are often psychoactive, but their effects are not well studied. It does appear that some sea cucumber toxins restrain the growth rate of tumour cells, which has sparked interest from cancer researchers.
The calcareous tests of echinoderms are used as a source of lime by farmers in areas where limestone is unavailable; indeed 4,000 tons of the animals are used annually for this purpose. This trade is often carried out in conjunction with shellfish farmers, for whom the starfish pose a major irritation by eating their stocks.
Echinoderms, like chordates, are deuterostomes and are therefore thought to be the most closely related of the major phyla to the chordates, being a sister group to chordates plus hemichordates. Chordates ( Phylum Chordata) are a group of Animals that includes the Vertebrates together with several closely related Invertebrates Deuterostomes (taxonomic term Deuterostomia; from the Greek "second mouth" are a superphylum of Animals They are a subtaxon of the Hemichordata is a phylum of worm-shaped marine Deuterostome Animals generally considered the sister group of the Echinoderms They date (Some believe that acorn worms are more closely related to echinoderms than chordates. The Acorn worms or Enteropneusta are a hemichordate class of Invertebrates Acorn worms are classified in the phylum Hemichordata, closely ) Because of a controversial interpretation of Homalozoa, a minority of classifiers place the echinoderms into the Chordata). Williamson (2003) disputes the links to hemichordates and chordates. They are based on larvae, which (Williamson claims) were later additions to life-histories. And pteropod hemichordates have larvae resembling trochophores, which would link them with annelids and molluscs. The phylogeny below is based on Smith (2005). It should be noted that this topology is not unilaterally agreed upon. Asteroids and Ophiuroids are frequently supported as sister groups by fossil and other molecular data.